1. CAS Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, PR China
2. Marine Ecology and Environmental Science Laboratory, Pilot National Laboratory for Marine Science and Technology (Qingdao), Qingdao 266200, PR China
3. University of Chinese Academy of Sciences, Beijing 100049, PR China
4. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao 266071, PR China
5. Jiaozhou Bay National Marine Ecosystem Research Station, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, PR China