1. Wildlife Biology Program, W.A. Franke College of Forestry & Conservation, University of Montana, 32 Campus Drive, Missoula, MT 59812, USA
2. School of Biological Sciences, Center for Reproductive Sciences, Washington State University, PO Box 644236, Pullman, WA 99164, USA
3. Department of Biology, Appalachian State University, 287 Rivers St, Boone, NC 28608, USA
4. School of the Environment, Washington State University, PO Box 644236, Pullman, WA 99163, USA
5. US Geological Survey, Northern Rocky Mountain Science Center, 32 Campus Drive, Missoula, MT 59812, USA