1. Department of Marine Ecology, GEOMAR Helmholtz Centre for Ocean Research Kiel, Düsternbrooker Weg 20, 24105 Kiel, Germany
2. Department of Biology and Geology, Faculty of Education, Alexandria University, 21526 Alexandria, Egypt
3. Environmental and Marine Biology, Åbo Akademi University, Artillerigatan 6, 20520 Åbo, Finland
4. Zoological Institute, Christian-Albrechts University, 24118 Kiel, Germany