1. Ecology, Evolution, and Behavior Program, Department of Plant Biology, Michigan State University, East Lansing, Michigan 48824, USA
2. Department of Anthropology, Michigan State University, East Lansing, Michigan 48824, USA
3. Michigan State University Museum, Michigan State University, East Lansing, Michigan 48824, USA
4. Department of Geography, Environment and Spatial Sciences, Michigan State University, East Lansing, Michigan 48824, USA
5. W. J. Beal Botanical Garden, Michigan State University, East Lansing, Michigan 48824, USA