1. School of Aquatic and Fishery Sciences, University of Washington, Seattle, WA 98115, USA
2. Environmental and Fisheries Sciences Division, Northwest Fisheries Science Center, National Marine Fisheries Service, National Oceanic & Atmospheric Administration, Seattle, WA 98112, USA
3. Washington Department of Fish and Wildlife, PO Box 43200, Olympia, WA 98504, USA
4. Conservation Biology Division, Northwest Fisheries Science Center, National Marine Fisheries Service, National Oceanic & Atmospheric Administration, Seattle, WA 98112, USA