1. Department of Biological Sciences, University of Manitoba, MB, R3T 2N2, Canada
2. Fisheries and Oceans Canada, Winnipeg, MB, R3T 2N6, Canada
3. Department of Oceanography, Dalhousie University, Halifax, NS, B3H 3Y1, Canada
4. Department of National Defence, Nanoose Bay, BC, V9P 9K8, Canada