1. Centre for Ocean and Atmospheric Sciences, School of Environmental Sciences, University of East Anglia, Norwich NR4 7TJ, UK
2. Centre for Environment, Fisheries & Aquaculture Science (Cefas), Lowestoft NR33 0HT, UK
3. Scottish Association for Marine Science, Scottish Marine Institute, Oban, Argyll PA37 1QA, UK
4. Department of Marine Sciences, University of Gothenburg (UGOT), 405 30 Gothenburg, Sweden
5. CNRS-Sorbonne Universités (UPMC Univ. Pierre et Marie Curie, Paris 06)-CNRS-IRD-MNHN, UMR 7159, Laboratoire d’Océanographie et de Climatologie (LOCEAN), Institut Pierre Simon Laplace (IPSL), Observatoire Ecce Terra, 75005 Paris, France