1. Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang, China; University of Chinese Academy of Sciences, Beijing, China
2. Research Assistant, Guangzhou Institute of Geography, Guangdong Academy of Sciences, Guangzhou, China; College of Earth and Environmental Sciences, Lanzhou University, Lanzhou, China
3. Professor, College of Geography Sciences, Qinghai Normal University, Qinghai, China; Academy of Plateau Science and Sustainability, Qinghai Normal University, Xining, China
4. Professor, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang, China; Academy of Plateau Science and Sustainability, Qinghai Normal University, Xining, China (corresponding author: )