1. Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, 100-Letiya Vladivostoku Ave, 159, Vladivostok 690022, Russia
2. Sobolev Institute of Geology and Mineralogy, Siberian Branch of the Russian Academy of Sciences, Academika Koptuga Ave., 3, Novosibirsk 630090, Russia; Laboratory of Mesozoic and Cenozoic Continental Ecosystems, Lenina Ave., 36, Tomsk State University, Tomsk 634050, Russia
3. Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, Saint Petersburg 199034, Russia
4. National Taiwan University, Roosevelt Road, Taipei 10617, Taiwan