1. Universidad Nacional San Antonio Abad del Cusco (UNSAAC), Cusco, Perú
2. Osa Conservation, Puerto Jiménez, Golfito, Costa Rica
3. Instituto Tecnológico de Costa Rica, Cartago, Costa Rica
4. Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, UK