1. Conservation Research Group, St. Albert's College, Kochi, 682 018, Kerala, India
2. Community Environmental Resource Center, Ashoka Trust for Research in Ecology and Environment, Alappuzha, Kerala, India
3. Department of Aquaculture, St. Albert's College, Kochi, 682 018, Kerala, India
4. Durrell Institute of Conservation and Ecology, University of Kent, Canterbury, CT2 7NR, Kent, UK Author for correspondence: