1. Institute of Biotechnology, National Dong Hwa University, Hualien, Taiwan, Republic of China
2. ‡Institute of Fisheries Sciences, National Taiwan University, Taipei, Taiwan 100, Republic of China
3. Institute of Zoology, Academia Sinica, Taipei, Taiwan 115, Republic of China
4. §Graduate Institute of Life Sciences, National Defense Medical Center, Taipei, Taiwan 100, Republic of China