1. Research Unit for Environmental Science and Management, Potchefstroom Campus, North-West University, Potchefstroom, South Africa
2. Protected Areas Research Group, Potchefstroom Campus, North-West University, South Africa
3. Biodiversity Research Centre, Namibian University of Science and Technology, Windhoek, Namibia
4. Namibia Nature Foundation, Windhoek, Namibia
5. School of Environmental Sciences, University of East Anglia, Norwich, UK