1. College of Land Science and Technology, China Agricultural University, Beijing, China
2. School of Environmental and Geographical Sciences and Wetlands Ecosystem Observation and Research Field Station, Shanghai Normal University, Shanghai, China
3. College of Land Science and Technology, China Agricultural University, Beijing, China; National Natural Science Foundation of China, Beijing, China
4. School of Environmental and Geographical Sciences and Wetlands Ecosystem Observation and Research Field Station, Shanghai Normal University,Shanghai, China