1. Nuffield Department of Orthopaedics Rheumatology and Musculoskeletal Sciences, The Botnar Research Centre, University of Oxford, Oxford, UK
2. NIHR Oxford Biomedical Research Centre, The Joint Research Office, Oxford, UK
3. Oxford University Hospitals NHS Foundation Trust Nuffield Orthopaedic Centre, Oxford, Oxfordshire, UK, London, UK
4. Dinwoodie Charitable Company and Royal College of Surgeons of England Research Fellow, London, UK
5. The MRC Weatherall Institute of Molecular Medicine, Oxford, UK