1. Department of Geography, Swansea University, School of Environment and Society, Singleton Park, Swansea SA2 8PP, UK,
2. Department of Geography, Swansea University, School of Environment and Society, Singleton Park, Swansea SA2 8PP, UK
3. Department of Physical Geography and Quaternary Geology, Stockholm University, SE-10691 Stockholm, Sweden, Abisko Scientific Research Station, the Royal Swedish Academy of Sciences, SE-98107 Abisko, Sweden
4. Finnish Forest Research Institute, Rovaniemi Research Unit, PO Box 16, FI-96301 Rovaniemi, Finland