1. Department of Ecology and Systematics, Division of Hydrobiology, University of Helsinki, PO Box 17 (Arkadiankatu 7), FIN-00014 Helsinki, Finland
2. Botanical Institute, University of Bergen, Allégaten 41, N-5007 Bergen, Norway, and Environmental Change Research Centre, University College London, 26 Bedford Way, London WC1H 0AP, UK
3. Geological Survey of Finland, PO Box 96, FIN-02151 Espoo, Finland
4. Department of Geography, PO Box 64, FIN-00014 University of Helsinki, Finland