1. Department of Geography, Loughborough University, Loughborough LE11 3TU, UK,
2. Environmental Change Research Centre, Department of Geography, Pearson Building, Gower Street, University College London, London WC1E 6BT, UK
3. Department of Geosciences, University of Massachusetts, Amherst MA 01003, USA
4. Institute of Marine Science, University of Alaska Fairbanks, Fairbanks AK 99775-7220, USA, Department of Biological Sciences, Idaho State University, Pocatello ID 83209-8007, USA