1. Alfred Wegener Institute for Polar and Marine Research, 14473 Potsdam, Germany,
2. Department of Biology & Bjerknes Centre for Climate Research, University of Bergen, Allégaten 41, N-5007 Bergen, Norway, and Environmental Change Research Centre, University College London, London WC1E 6BT, UK
3. Alfred Wegener Institute for Polar and Marine Research, 14473 Potsdam, Germany, State Key Laboratory of Vegetation and Environmental Change, Institute of Botany, Chinese Academy of Sciences, 100093 Beijing, China
4. MOE Key Laboratory of Western China's Environmental System, College of Earth and Environmental Sciences, Lanzhou University, 730000 Lanzhou, China
5. Department of Ecology, College of Environmental Sciences, Beijing University, 100871 Beijing, China
6. Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences, 210008 Nanjing, China
7. Geophysical Institute, University of Alaska Fairbanks, 903 Koyukuk Drive, Fairbanks, AK 99775, USA