1. *Laboratory of Fish Molecular Immunology, College of Fisheries and Life Science, Shanghai Ocean University, Shanghai, China
2. †Laboratory of Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao, China
3. ‡National Pathogen Collection Center for Aquatic Animals, Shanghai Ocean University, Shanghai, China
4. §Key Laboratory of Freshwater Aquatic Genetic Resources, Ministry of Agriculture and Rural Affairs, Shanghai Ocean University, Shanghai, China
5. ¶Marine Biomedical Science and Technology Innovation Platform of Lin-gang Special Area, Shanghai, China