1. *Laboratory of Immune Regulation, Department of Virus Research, Institute for Life and Medical Sciences, Kyoto University, Kyoto, Japan;
2. †Graduate School of Biostudies, Kyoto University, Kyoto, Japan;
3. ‡Department of Human Health Sciences, Graduate School of Medicine, Kyoto University, Kyoto, Japan;
4. §Department of Respiratory Medicine, Graduate School of Medicine, Kyoto University, Kyoto, Japan;
5. ¶Department of Medical Chemistry, Graduate School of Medicine, Kyoto University, Kyoto, Japan;
6. ‖Graduate School of Life Science and Engineering, Iryo Sosei University, Iwaki, Japan;
7. #Research Institute of Innovative Medicine, Tokiwa Foundation, Iwaki, Japan; and
8. **School of Medicine, Fukushima Medical University, Fukushima, Japan