1. Kansai Research Center, Forestry and Forest Products Research Institute, 68 Nagaikyutaro, Momoyama, Fushimi, Kyoto, 612-0855 Japan
2. Forest Research Institute Malaysia, 52109 Kepong, Selangor Darul Ehsan, Malaysia
3. Makino Herbarium, Tokyo Metropolitan University, Minamiosawa, Hachioji, Tokyo, 192-0397, Japan