1. Biology Program, Department of Science, Faculty of Science and Technology, Prince of Songkla University (Pattani)
2. Biology Program, Faculty of Science, Ubon Ratchathani Rajabhat University
3. Applied Taxonomic Research Center (ATRC), Department of Biology, Faculty of Science, Khon Kaen University
4. Biology Program, Faculty of Science, Buriram Rajabhat University