1. Arano, H. 1965. The karyotypes and the speciations in the subfamily Carduoideae (Compositae) of Japan xviii. Jpn. J. Bot. 19: 31–67.
2. Arohonka, T. 1982. Chromosome counts of vascular plants of the island Seili in Nauvo, SW Finland. Turunyliopiston biologian–laitoskan julkaisuja 3: 1–12.
3. Aswal, B. S. and Mehrotra, B. N. 1994. Flora of Lahaul-Spiti (a cold desert in Northwest Himalaya). Bishen Singh Mahendra Pal Singh, Dehradun.
4. Bala, S. and Gupta, R. C. 2011. In: Marhold, K. (ed.). IAPT/IOPB Chromosome data. 12. Taxon 60: 1784–1796.
5. Bala, S. and Gupta, R. C. 2013. Male meiosis and chromosome number in Asteraceae family from district Kangra of H.P. (Western Himalaya). Intl. J. Bot. Res. 3: 43–58.