1. 1 Department of Organismic and Evolutionary Biology, Harvard Herbarium, 22 Divinity Avenue, Cambridge, MA 02138, U.S.A.
2. 2 Blaihofstr. 42, 72074 Tübingen, Germany.
3. 3 Department of Botany, Ecology and Plant Physiology, University of La Laguna, 38200 Tenerife, Canary Islands, Spain.