1. College of Forestry and Landscape Architecture, Guangdong Key Laboratory for Innovative Development and Utilization of Forest Plant Germplasm, South China Agricultural University, Guangzhou 510642, China
2. Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin 541006, China.
3. South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China