1. Kyoto University, Graduate School of Engineering, Kyoto 615-8540, Japan.(corresponding author).
2. University of Hyogo, Graduate School of Life Science, Hyogo 671-2280, Japan..
3. Japan Agency for Marine-Earth Science and Technology, Kanagawa 237-0061, Japan..
4. Nagoya University, Graduate School of Environmental Studies, Aichi 464-8603, Japan..