1. Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560; ORCID: 0000-0002-2875-0474; Email: .
2. Sam Noble Oklahoma Museum of Natural History and Department of Biology, University of Oklahoma, Norman, Oklahoma 73072; ORCID: 0000-0002-5126-1345; Email:
3. Smithsonian Tropical Research Institute, Balboa, Republic of Panama; ORCID: 0000-0003-3972-149X; Email: .
4. University of Washington, School of Aquatic and Fishery Sciences, Burke Museum of Natural History and Culture, Seattle, Washington 98195; Email: .