1. Department of Biology and Ecology Center, Utah State University, Logan, Utah 84322-5305; Email: (MK) ; (SBH) spencerbrucehudson@gmail.com; (EEV) emilyevirgin@gmail.com; (HBP) plylar@aggiemail.usu.edu; (SSF) Susannah.french@usu.edu; and (AHS) savitzky