1. Department of Biological Sciences, Lock Haven University, 301 W Church Street, Lock Haven, Pennsylvania 17745; Email: (HNR) ; and (JD) jmd240@lockhaven.edu. Send reprint requests to JD.
2. Department of Ecology, Evolution, and Organismal Biology, Kennesaw State University, 370 Paulding Ave NW, Kennesaw, Georgia 30144; Email: .