1. Center for Molecular Biodiversity Research, National Museum of Nature and Science, 4-1-1 Amakubo, Tsukuba, Ibaraki 305-0005, Japan; Email: . Send reprint requests to this address.
2. School of Biological Sciences, Universiti Sains Malaysia, 11800, Penang, Malaysia; Email: .
3. The Kagoshima University Museum, 1-21-30 Korimoto, Kagoshima 890-0065, Japan; Email: .