1. Institut des Sciences de l’Evolution de Montpellier (ISEM), CNRS, IRD, EPHE, Université de Montpellier, Montpellier, France
2. National Museum and Centre for Environmental Management, University of the Free State, Bloemfontein, South Africa
3. Department of Natural Sciences, National Museums Scotland, Edinburgh, United Kingdom
4. CNRS, Sorbonne Université, CNRS, ABiMS, Station Biologique de Roscoff, Roscoff, France
5. Institut Universitaire de France (IUF), Paris, France