1. Department of Cell Biology, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands
2. Department of Molecular Pharmacology, Groningen Research Institute of Pharmacy (GRIP), Faculty of Science and Engineering, University of Groningen, Groningen, The Netherlands
3. Wellcome Trust Centre for Human Genetics, Oxford, United Kingdom
4. Office of the President, Tufts University, Medford, United States