1. European Molecular Biology Laboratory, Heidelberg, Germany
2. School of Life and Environmental Sciences, University of Sydney, Sydney, Australia
3. Computational Biology and Biological Physics, Department of Astronomy and Theoretical Physics, Lund University, Lund, Sweden
4. Division of Biology and Biological Engineering, California Institute of Technology, Howard Hughes Medical Institute, Pasadena, United States
5. Sainsbury Laboratory, University of Cambridge, Cambridge, United Kingdom
6. Department of Applied Mathematics and Theoretical Physics, University of Cambridge, Cambridge, United Kingdom