1. Division of Biological Science, Graduate School of Science, Nagoya University, Nagoya, Japan
2. Division of Molecular Cell Engineering, National Institute of Genetics, Research Organization of Information and Systems, Shizuoka, Japan
3. Department of Genetics, SOKENDAI, Shizuoka, Japan
4. Precursory Research for Embryonic Science and Technology (PRESTO) Program, Japan Science and Technology Agency, Saitama, Japan