1. INRAE, UMR1391 ISPA,Villenave d’Ornon,France,33140
2. University of Toulouse, National Center for Scientific Reaseach (CNRS), Institute for Research and Development (IRD), UPS. Observatory Midi-Pyrénées (OMP),Geosciences Environment Toulouse (GET),Toulouse,France,31400
3. University of Montpellier,ESPACE-DEV, IRD,Montpellier,France,34093