1. Department of Biological Sciences, University of Cincinnati, Cincinnati, Ohio 45221-0006, U.S.A.;, Email: eric.tepe@uc.edu
2. Department of Ecology and Evolutionary Biology, University of Connecticut, Storrs, Connecticut 06269-3043, U.S.A.
3. USDA Agricultural Research Service, Department of Horticulture, University of Wisconsin, Madison, Wisconsin 53706-1590, U.S.A.
4. Biology Department, University of Utah, Salt Lake City, Utah 84112, U.S.A.