1. Missouri Botanical Garden, P.O. Box 299, St. Louis, Missouri 63166-0299, U.S.A, Contributed equally to this work;, Email: ortiz-gentry@mobot.org
2. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, P.R. China, Contributed equally to this work;, Email: wangwei1127@ibcas.ac.cn
3. Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla 666303, P.R. China, Current address: 11-13 rue Garnier, 92200 Neuilly sur Seine, France,
Contributed equally to this work
4. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, P.R. China