1. Missouri Botanical Garden, P.O. Box 299, St. Louis, Missouri 63166-0299, U.S.A.;, Email: ihsan.al-shehbaz@mobot.org
2. Department of Biodiversity and Plant Systematics, Centre for Organismal Studies, Heidelberg University, Im Neuenheimer Feld 345, 69120 Heidelberg, Germany; and South-Siberian Botanical Garden, Altai State University, Lenin str.
61, 656049 Barnaul, Russia