1. Department of Botany, P.O. Box 77000, Nelson Mandela Metropolitan University, Port Elizabeth, 6031 South Africa, Centre for Functional Ecology, Departamento de Ci�ncias da Vida, Universidade de Coimbra, 3001-455 Coimbra,
Portugal;, Email: smithgideon1@gmail.com
2. Department of Botany, P.O. Box 77000, Nelson Mandela Metropolitan University, Port Elizabeth, 6031 South Africa, Centre for Functional Ecology, Departamento de Ci�ncias da Vida, Universidade de Coimbra, 3001-455 Coimbra, Portugal
3. National Plant Data Team, East National Technology Support Center, Natural Resources Conservation Service, United States Department of Agriculture, 2901 E Lee Street, Greensboro, North Carolina, U.S.A