1. School of Biological Sciences, The University of Hong Kong, Hong Kong, China
2. Biosystematics group, Wageningen University, Droevendaalsesteeg 1, 6708 PB Wageningen, The Netherlands, Naturalis Biodiversity Center (Section NHN), Herbarium Vadense, Darwinweg 2, 2333 CR Leiden, The Netherlands;, Email: Paul.Hoekstra@naturalis.nl
3. School of Biological Sciences, The University of Hong Kong, Hong Kong, China, School of Science and Technology, The Open University of Hong Kong, Ho Man Tin, Kowloon, Hong Kong, China
4. School of Biological Sciences, The University of Hong Kong, Hong Kong, China, Singapore Botanic Gardens, 1 Cluny Road, Singapore 259569, Singapore
5. Biosystematics group, Wageningen University, Droevendaalsesteeg 1, 6708 PB Wageningen, The Netherlands, Naturalis Biodiversity Center (Section NHN), Herbarium Vadense, Darwinweg 2, 2333 CR Leiden, The Netherlands
6. Biosystematics group, Wageningen University, Droevendaalsesteeg 1, 6708 PB Wageningen, The Netherlands
7. School of Biological Sciences, The University of Hong Kong, Hong Kong, China;, Email: saunders@hku.hk