1. Utrecht University, Department of Biology, Padualaan 8, 3584CH Utrecht, the Netherlands, Department of Life Sciences, Natural History Museum, Cromwell Road, SW7 5BD London, U.K.
2. 390 chemin des Vignes vieilles, 84120 Mirabeau, France
3. Institute of Systematic Botany, The New York Botanical Garden, Bronx, New York 10458-5126, U.S.A.
4. Department of Life Sciences, Natural History Museum, Cromwell Road, SW7 5BD London, U.K., CSIRO, Australian National Herbarium, P.O. Box 1600, Canberra, ACT, 2601, Australia;, Email: Cecile.Gueidan@csiro.au