1. Department of Animal Nutrition and Feed Science, College of Animal Science and Technology, Huazhong Agricultural University, Wuhan 430070, P. R. China
2. Key Lab of Agricultural Animal Genetics, Breeding and Reproduction of Ministry of Education & Key Lab of Swine Genetics and Breeding of Ministry of Agriculture, Huazhong Agricultural University, Wuhan 430070, P. R. China
3. YangXiang Joint Stock Company, Guigang 537000, P. R. China
4. The Cooperative Innovation Center for Sustainable Pig Production, Wuhan 430070, P. R. China