1. Department of Agricultural, Food and Nutritional Science, University of Alberta, Edmonton T6G 2C8, Canada
2. Alberta Agriculture and Rural Development, Edmonton T6H 5T6, Canadda
3. Alberta Agriculture and Rural Development, Lacombe Research Centre, Lacombe, Alberta T4L 1W1, Canada
4. Beefbooster Inc., Calgary, Alberta T1Y 5E4, Canada
5. Center for Genetic Improvement of Livestock, University of Guelph, Guelph, Ontario N1G 2W1, Canada