1. Victorian Department of Environment and Primary Industries, Bundoora, VIC 3083, Australia
2. New South Wales Department of Primary Industries, Beef Industry Centre, University of New England, Armidale, NSW 2351, Australia
3. School of Veterinary Science, University of Melbourne, Parkville, VIC 3010, Australia
4. Murdoch Children's Research Institute, Royal Children's Hospital, Parkville, VIC 3052, Australia