1. Clean Energy Research Center, Korea Institute of Science and Technology (KIST), Seoul 02792, Republic of Korea
2. Laboratory of Marine Microbiology, Division of Convergence on Marine Science, Korea Maritime and Ocean University, Busan 49112, Republic of Korea
3. Department of Marine Bioscience and Environment, Korea Maritime & Ocean University, Busan 49112, Republic of Korea
4. National Institute of Biological Resources, Environmental Research Complex, Incheon 22689, Republic of Korea