1. Animal Biotechnology Division, National Institute of Animal Science, Rural Development Administration, Wanjugun, Jeollabuk-do 55365, Republic of Korea
2. Animal Genomics and Bioinformatics Division, National Institute of Animal Science, Rural Development Administration, Wanju-gun, Jeollabuk-do 55365, Republic of Korea
3. Hanwoo Research Institute, National Institute of Animal Science, Rural Development Administration, Wanju-gun, Jeollabuk-do 55365, Republic of Korea