1. Department of Biology, Eastern Michigan University, Ypsilanti, Michigan, 48197, USA
2. Herpetological Resource and Management, PO Box 110, Chelsea, Michigan, 48118, USA
3. Division of Ecological and Water Resources, Minnesota Department of Natural Resources, Saint Paul, Minnesota, 55155, USA
4. California Department of Fish and Wildlife, North Central Region, 1701 Nimbus Rd., Rancho Cordova, California, 95670, USA
5. US Geological Survey, Vermont Cooperative Fish and Wildlife Research Unit, Rubenstein School of Environment and Natural Resources, University of Vermont, Burlington, Vermont, 05405, USA
6. Department of Biology, University of Vermont, Burlington, Vermont, 05405, USA