1. Museu de História Natural e da Ciência da Universidade do Porto, Praça de Gomes Teixeira, 4099-002 Porto, Portugal
2. Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania, 19085-1699, USA
3. Laboratoire d'Hérpétologie, Departement de Biologie, Centre de Recherche en Sciences Naturelles, Lwiro, Democratic Republic of the Congo
4. Department of Biological Sciences, University of Texas at El Paso, 500 W. University Avenue, El Paso, Texas, 79912, USA