1. Systematics Lab, Department of Environmental Studies, University of Delhi, Delhi 110007, India; e-mail: , sdbiju@fas.harvard.edu.
2. Andaman and Nicobar Regional Centre, Zoological Survey of India, Port Blair 744102, Andaman and Nicobar Islands, India.
3. Department of Ecology, Evolution, and Behavior, University of Minnesota–Twin Cities, St. Paul, Minnesota 55108, U.S.A.
4. Museum of Comparative Zoology and Department of Organismic and Evolutionary Biology, Harvard University, 26 Oxford Street, Cambridge, Massachusetts 02138, U.S.A.; e-mail: .